Marc-André Villard 
Assistant Professor

Marc-Andre VillardI am a terrestrial ecologist with a particular interest for species response to habitat fragmentation and degradation by human activities.  I frequently use forest songbirds as a model to study the effects of landscape structure on reproductive success, functional connectivity, and other processes influencing the viability of their populations. My students, colleagues and I are also investigating the effects of climate change and shoreline protection on the staging ecology of shorebirds.

This research is funded by NSERC, NBIF, and NBWTF.


Fall 2018: Introductory Ecology (BIOL 2101), Animal Behaviour (BIOL 3401)

Winter 2019: Ornithology (BIOL 3651), and Conservation Biology (BIOL 3811). 

I strive to illustrate the concepts seen in class through examples taken from recent publications or phenomena that can be readily observed in the field.




Postdoctoral Researcher – Alberta (1994)

Postdoctoral Researcher – Brigham Young (1993)

Ph.D. – Carleton (1991)

B.Sc. – Montréal (1986)



Themes of current projects:

Effects of intensive forest management on functional connectivity of songbird populations

Insectivorous songbirds as early indicators of spruce budworm outbreaks

Predictors of nest predation risk in forest songbirds and woodpeckers

Effects of coastal storms on shorebird staging ecology


Selected Publications (*graduate students)

*Torrenta, R., Lacoste, F., and Villard, M.-A.  In press.  Loss and fragmentation of mature woodland reduce the habitat niche breadth of forest birds. Landscape Ecology. 

Villard, M.-A. and Foppen, R.  2018.  Ecological adaptations of birds to forest environments. Pp. 51-78 in Mikusinski, G. et al. (editors). Ecology and Conservation of Forest Birds. Cambridge University Press.

*Fiola, M.-L., *Vernouillet, A., and Villard, M.-A.  2017.  Linking songbird nest predation to seedling density: sugar maple masting as a resource pulse in a forest food web. Ecology and Evolution 7(24): 10733-10742.

*Torrenta, R. and Villard, M.-A.  2017.  A test of the habitat amount hypothesis as an explanation for the species richness of forest bird assemblages.  Journal of Biogeography 44: 1791-1801.

*MacKay, A., Allard, M., and Villard, M.-A. 2014. Capacity of older plantations to host bird assemblages of naturally-regenerated conifer forests: a test at stand and landscape levels.  Biological Conservation 170: 110-119.

Villard, M.-A. and Metzger, J.P.  2014.  Beyond the fragmentation debate: a conceptual model to predict when habitat configuration really matters.  Journal of Applied Ecology 51: 309-318.

*Haché, S., Bayne, E.M., and Villard, M.-A.  2013.  Evidence for an ideal free distribution in a breeding population of a territorial songbird.  Ecology 94: 861-869.

Villard, M.-A. and *Haché, S.  2012.  Conifer plantations consistently act as barriers to movements in a deciduous forest bird: a translocation experiment.  Biological Conservation 155: 33-37

*Haché, S. and Villard, M.-A.  2010.  Age-specific response of a migratory bird to an experimental alteration of its habitat.  Journal of Animal Ecology 79: 897-905.

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Contact information 

Office : Flem 207

Phone : 506-364-2522

Email :

Office hours : Monday 1:30-5:00; Thursday 1:30-5:00